Bangkok, September 16: More than half of the 147 tigers confiscated from a controversial Thai temple have died, park officials said Monday, blaming genetic problems linked to in-breeding at the once money-spinning tourist attraction.

For years, the Wat Pha Luang Ta Bua temple in the western province of Kanchanaburi pulled tourist hordes who could be photographed -- for a fee -- next to scores of tigers.

But in 2016 park officials began a lengthy operation to remove the big cats amid allegations of mismanagement, and claims the creatures were being exploited.

Dozens of dead cubs were found kept in freezers, sparking claims the carcasses were being sold by a temple rumoured to have raked in hundreds of thousands of dollars each year from visitors.

Tiger parts can fetch enormous sums in China and Vietnam, where they are falsely believed to have medicinal properties.

The surviving adults were taken to two breeding stations in nearby Ratchaburi province but only 61 of the 147 have survived so far, parks officials told reporters.

"It could be linked to in-breeding," Pattarapol Maneeon, of the Department of National Parks, Wildlife and Plant Conservation said.

"They had genetic problems which posed risks to body and immune system."

Many suffered from tongue paralysis, breathing problems and lack of appetite that led to fatal seizures.

- Legal cases ongoing -

"Most of the tigers were already in distressed state stemming from the transportation and change of location... later their health problems emerged," Sunthorn Chaiwattana, another department official said.

Legal cases against the temple are ongoing.

Conservationists questioned whether authorities had looked after the seized animals appropriately, with small, cramped cages enabling the spread of disease.

"To be very honest who would be ready to take in so many tigers at once?" said Edwin Wiek, founder of the Wildlife Friends Foundation Thailand.

Conditions at the enclosures were "not good enough to house so many tigers and the set up was wrong", he added.

Tens of millions visit Thailand every year, and a lucrative wildlife tourism industry has grown in-step with visitor numbers.

But critics say cash often trumps animal welfare at many attractions.

For a price, visitors can ride and bathe with elephants, hold monkeys and pose for selfies with tigers.

Animal rights groups have long criticised the industry, with chained up animals kept in small quarters with inadequate veterinary care, or forced to perform tricks for tourists.

(The above story first appeared on LatestLY on Sep 16, 2019 04:14 PM IST. For more news and updates on politics, world, sports, entertainment and lifestyle, log on to our website latestly.com).